Gymnognathus Schoenherr, 1826: 37

Species and distribution according to Rheinheimer (2004, 2021). Additional distribution records according to Jordan (1906), Wolfrum (1929), Blackwelder (1947), Perger & Guerra (2016), Orellana & Barrios (2021).

Type species. Anthribus ancora Germar, 1824: 177

Synonyms. Gymnognathus Schoenherr, 1823: 1135; Analotes Schoenherr, 1839: 198

Diversity. 96 described species

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Table of Contents

  1. Gymnognathus abundans Jordan, 1906: 339
  2. Gymnognathus acastus Jordan, 1937: 215
  3. Gymnognathus acutangulus Jordan, 1895: 170
  4. Gymnognathus ada Jordan, 1904: 243
  5. Gymnognathus affinis Jordan, 1895: 251
  6. Gymnognathus alma Jordan, 1904: 247
  7. Gymnognathus ampulla Jordan, 1906: 338
  8. Gymnognathus ancora (Germar, 1824: 177)
  9. Gymnognathus aulicus Jordan, 1937: 227
  10. Gymnognathus barclayi Perger & Guerra, 2016: 279
  11. Gymnognathus bella Jordan, 1904: 245
  12. Gymnognathus bifasciatus Rheinheimer, 2021: 308
  13. Gymnognathus biloris Jordan, 1937: 227
  14. Gymnognathus blanca Jordan, 1904: 243
  15. Gymnognathus bohlsi Jordan, 1895: 172
  16. Gymnognathus brevirostris Jordan, 1895: 250
  17. Gymnognathus bryanti Jordan, 1904: 213
  18. Gymnognathus calus Jordan, 1906: 339
  19. Gymnognathus cerdai Rheinheimer, 2021: 309
  20. Gymnognathus chiricahuae Sleeper, 1954: 118
  21. Gymnognathus cincticollis Jordan, 1895: 168
  22. Gymnognathus clara Jordan, 1904: 245
  23. Gymnognathus clathratus Jordan, 1895: 170
  24. Gymnognathus claudia Jordan, 1906: 337
  25. Gymnognathus clelia Jordan, 1904: 248
  26. Gymnognathus compar Jordan, 1895: 167
  27. Gymnognathus comptus Jordan, 1904: 252
  28. Gymnognathus cordiger Frieser, 1959: 418
  29. Gymnognathus coronatus Jordan, 1904: 246
  30. Gymnognathus dagmari Frieser, 1981: 208
  31. Gymnognathus daguanus Jordan, 1897: 360
  32. Gymnognathus decorus Perroud, 1853: 402
  33. Gymnognathus depressibasis Rheinheimer, 2021: 311
  34. Gymnognathus discoideus (Fahraeus, 1839: 199)
  35. Gymnognathus doris Jordan, 1937: 228
  36. Gymnognathus dorsonotatus Fahraeus, 1839: 202
  37. Gymnognathus editha Jordan, 1904: 250
  38. Gymnognathus elisa Jordan, 1937: 231
  39. Gymnognathus emma Jordan, 1904: 244
  40. Gymnognathus erna Jordan, 1904: 248
  41. Gymnognathus extensus Jordan, 1904: 242
  42. Gymnognathus fahraei Fahraeus, 1939: 200
  43. Gymnognathus flexuosus Jordan, 1895: 173
  44. Gymnognathus germaini Jordan, 1897: 173
  45. Gymnognathus hamatus Jordan, 1904: 251
  46. Gymnognathus hedys Jordan, 1937: 216
  47. Gymnognathus helena Jordan, 1904: 249
  48. Gymnognathus helma Jordan, 1937: 213
  49. Gymnognathus hetarus Jordan, 1937: 213
  50. Gymnognathus hilda Jordan, 1904: 244
  51. Gymnognathus inca Jordan, 1937: 218
  52. Gymnognathus indagatus Wolfrum, 1931: 71
  53. Gymnognathus iphis Jordan, 1937: 221
  54. Gymnognathus iris Jordan, 1937: 216
  55. Gymnognathus irma Jordan, 1904: 247
  56. Gymnognathus leucomelas Jordan, 1904: 251
  57. Gymnognathus libussa Jordan, 1937: 220
  58. Gymnognathus lotus Jordan, 1937: 224
  59. Gymnognathus lusia Jordan, 1937: 219
  60. Gymnognathus lyrestes Jordan, 1937: 214
  61. Gymnognathus mariana Jordan, 1904: 243
  62. Gymnognathus martha Jordan, 1904: 250
  63. Gymnognathus mexicanus Jordan, 1906: 338
  64. Gymnognathus molitor Jordan, 1895: 249
  65. Gymnognathus mollis Jordan, 1937: 217
  66. Gymnognathus moranus Jordan, 1937: 222
  67. Gymnognathus nanus Jordan, 1904: 248
  68. Gymnognathus nebulosus Motschulsky, 1875: 235
  69. Gymnognathus nica Jordan, 1937: 222
  70. Gymnognathus nubilus Jordan, 1904: 246
  71. Gymnognathus ophiopis Dalman, 1833: 163
  72. Gymnognathus ornatus Jordan, 1895: 171
  73. Gymnognathus pentilus Jordan, 1937: 225
  74. Gymnognathus phanerus Jordan, 1937: 229
  75. Gymnognathus pindonis Jordan, 1937: 231
  76. Gymnognathus polius Jordan, 1906: 337
  77. Gymnognathus procerus Jordan, 1937: 226
  78. Gymnognathus pulchellus Jordan, 1937: 229
  79. Gymnognathus pulcher Jordan, 1906: 340
  80. Gymnognathus regalis Jordan, 1937: 225
  81. Gymnognathus robustus Jordan, 1895: 169
  82. Gymnognathus ruficlava Jordan, 1906: 339
  83. Gymnognathus scalaris Jordan, 1906: 336
  84. Gymnognathus scolytinus Jordan, 1904: 252
  85. Gymnognathus signatus Gyllenhal, 1833: 165
  86. Gymnognathus soror Jordan, 1904: 249
  87. Gymnognathus stigmenus Wolfrum, 1955: 676
  88. Gymnognathus talis Jordan, 1937: 230
  89. Gymnognathus tenuis Jordan, 1895: 250
  90. Gymnognathus thecla Jordan, 1906: 338
  91. Gymnognathus triangularis Valentine, 1998: 269
  92. Gymnognathus uta Wolfrum, 1955: 676
  93. Gymnognathus vanda Jordan, 1937: 219
  94. Gymnognathus variicornis Jordan, 1895: 174
  95. Gymnognathus vicinus Jekel, 1855: 117
  96. Gymnognathus viticollis Jordan, 1897: 175
  97. Gymnognathus groups according to Jordan (1937)

Gymnognathus abundans Jordan, 1906: 339

Distribution. Guatemala [type].

Gymnognathus abundans Gymnognathus abundans
Gymnognathus abundans Jordan, 1.) lateral view, 2.) dorsal view. Specimens deposited at Arizona State University Biocollections (ASUCOB, ASUHIC). Images and data available at ecdysis portal and GBIF. Images: Samanta Orellana.

Gymnognathus acastus Jordan, 1937: 215

Distribution. French Guiana [type].

Gymnognathus acutangulus Jordan, 1895: 170

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2016: 314.

Gymnognathus ada Jordan, 1904: 243

Distribution. Brazil [type].

Gymnognathus affinis Jordan, 1895: 251

Distribution. Brazil [type].

Gymnognathus alma Jordan, 1904: 247

Distribution. Brazil [type].

Gymnognathus ampulla Jordan, 1906: 338

Distribution. Panama [type].

Gymnognathus ancora (Germar, 1824: 177)

Type species

Distribution. Brazil [type].

Gymnognathus aulicus Jordan, 1937: 227

Distribution. Brazil [type].

Gymnognathus barclayi Perger & Guerra, 2016: 279

Distribution. Brazil [type].

Images. Perger & Guerra, 2016: 280.

Gymnognathus bella Jordan, 1904: 245

Distribution. Brazil [type].

Gymnognathus bifasciatus Rheinheimer, 2021: 308

Distribution. French Guiana [type].

Images. Rheinheimer, 2016: 313.

Gymnognathus biloris Jordan, 1937: 227

Distribution. Brazil [type].

Gymnognathus blanca Jordan, 1904: 243

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2016: 315.

Gymnognathus bohlsi Jordan, 1895: 172

Distribution. Paraguay [type].

Images. Rheinheimer, 2004: 239.

Gymnognathus brevirostris Jordan, 1895: 250

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2016: 317.

Gymnognathus bryanti Jordan, 1904: 213

Distribution. Brazil [type].

Gymnognathus calus Jordan, 1906: 339

Distribution. Panama [type].

Gymnognathus ruficlava
Gymnognathus calus Jordan: 1.) dorsal view. Illustration in Jordan 1906, digitized by the Smithsonian Libraries under a CC0-Public Domain License. Modified by Samanta Orellana.

Gymnognathus cerdai Rheinheimer, 2021: 309

Distribution. French Guiana [type].

Images. Rheinheimer, 2016: 315.

Gymnognathus chiricahuae Sleeper, 1954: 118

Distribution. United States of America [type].

Gymnognathus cincticollis Jordan, 1895: 168

Distribution. Brazil [type].

Gymnognathus clara Jordan, 1904: 245

Distribution. Brazil [type].

Gymnognathus clathratus Jordan, 1895: 170

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2004: 240; Rheinheimer, 2016: 313.

Gymnognathus claudia Jordan, 1906: 337

Distribution. Nicaragua [type], Panama.

Gymnognathus claudia Gymnognathus claudia
Gymnognathus claudia Jordan, 1.) lateral view, 2.) dorsal view.. Specimens deposited at Arizona State University Biocollections (ASUCOB, ASUHIC). Images and data available at ecdysis portal and GBIF. Images: Samanta Orellana.

Gymnognathus clelia Jordan, 1904: 248

Distribution. Brazil [type].

Gymnognathus compar Jordan, 1895: 167

Distribution. Brazil [type].

Gymnognathus comptus Jordan, 1904: 252

Distribution. Brazil [type].

Gymnognathus cordiger Frieser, 1959: 418

Distribution. Bolivia [type].

Images. Rheinheimer, 2004: 240.

Gymnognathus coronatus Jordan, 1904: 246

Distribution. Panama, Brazil [type].

Gymnognathus dagmari Frieser, 1981: 208

Distribution. Brazil [type].

Images. Rheinheimer, 2004: 240.

Gymnognathus daguanus Jordan, 1897: 360

Distribution. Panama, Colombia [type].

Gymnognathus daguanus Gymnognathus daguanus
Gymnognathus daguanus Jordan: 1.) lateral view, 2.) dorsal view. Specimen deposited at Arizona State University Biocollections (ASUCOB, ASUHIC). Images and data available at ecdysis portal and GBIF. Images: Samanta Orellana.

Gymnognathus decorus Perroud, 1853: 402

Distribution. Brazil [type].

Gymnognathus depressibasis Rheinheimer, 2021: 311

Distribution. French Guiana [type].

Images. Rheinheimer, 2016: 317.

Gymnognathus discoideus (Fahraeus, 1839: 199)

Distribution. Brazil [type], Paraguay.

Gymnognathus doris Jordan, 1937: 228

Distribution. Brazil [type].

Images. Rheinheimer, 2016: 313.

Gymnognathus dorsonotatus Fahraeus, 1839: 202

Distribution. Brazil [type].

Gymnognathus editha Jordan, 1904: 250

Distribution. Brazil [type].

Gymnognathus elisa Jordan, 1937: 231

Distribution. Brazil [type].

Gymnognathus emma Jordan, 1904: 244

Distribution. Brazil [type].

Gymnognathus erna Jordan, 1904: 248

Distribution. Brazil [type].

Gymnognathus extensus Jordan, 1904: 242

Distribution. Brazil [type].

Gymnognathus fahraei Fahraeus, 1939: 200

Distribution. Brazil [type].

Gymnognathus flexuosus Jordan, 1895: 173

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2016: 316.

Gymnognathus germaini Jordan, 1897: 173

Distribution. Bolivia [type].

Gymnognathus hamatus Jordan, 1904: 251

Distribution. Brazil [type].

Gymnognathus hedys Jordan, 1937: 216

Distribution. Panama [type].

Gymnognathus hedys
Gymnognathus hedys Jordan: 1.) lateral view. Illustration in Jordan 1906, digitized by the Smithsonian Libraries under a CC0-Public Domain License. Modified by Samanta Orellana.

Gymnognathus helena Jordan, 1904: 249

Distribution. Colombia [type].

Gymnognathus helma Jordan, 1937: 213

Distribution. Brazil [type].

Gymnognathus hetarus Jordan, 1937: 213

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2004: 240; Rheinheimer 2016: 313.

Gymnognathus hilda Jordan, 1904: 244

Distribution. Brazil [type].

Gymnognathus inca Jordan, 1937: 218

Distribution. Peru [type].

Gymnognathus indagatus Wolfrum, 1931: 71

Distribution. Costa Rica [type].

Gymnognathus iphis Jordan, 1937: 221

Distribution. Costa Rica [type].

Gymnognathus iris Jordan, 1937: 216

Distribution. Brazil, French Guiana.

Images. Rheinheimer, 2016: 315.

Gymnognathus irma Jordan, 1904: 247

Distribution. Brazil [type].

Gymnognathus leucomelas Jordan, 1904: 251

Distribution. Brazil [type].

Gymnognathus libussa Jordan, 1937: 220

Distribution. French Guiana [type].

Images. Rheinheimer, 2016: 316.

Gymnognathus lotus Jordan, 1937: 224

Distribution. Brazil [type], Bolivia, French Guiana.

Images. Rheinheimer, 2016: 314.

Gymnognathus lotus
Gymnognathus lotus Jordan, 1.) lateral view. Specimens deposited at Arizona State University Biocollections (ASUCOB, ASUHIC). Images and data available at ecdysis portal and GBIF. Images: Samanta Orellana.

Gymnognathus lusia Jordan, 1937: 219

Distribution. French Guiana [type].

Images. Rheinheimer, 2016: 315.

Gymnognathus lyrestes Jordan, 1937: 214

Distribution. Panama, Brazil [type], French Guiana.

Images. Rheinheimer, 2016: 313, Orellana & Barrios, 2021: 30.

Gymnognathus mariana Jordan, 1904: 243

Distribution. Brazil [type].

Gymnognathus martha Jordan, 1904: 250

Distribution. Brazil [type].

Gymnognathus mexicanus Jordan, 1906: 338

Distribution. Mexico [type].

Gymnognathus mexicanus
Gymnognathus mexicanus Jordan: 1.) dorsal view. Illustration in Jordan 1906, digitized by the Smithsonian Libraries under a CC0-Public Domain License. Modified by Samanta Orellana.

Gymnognathus molitor Jordan, 1895: 249

Distribution. Guyana [type].

Images. Rheinheimer, 2016: 313.

Gymnognathus mollis Jordan, 1937: 217

Distribution. Brazil [type], French Guiana.

Images. Rheinheimer, 2004: 240; Rheinheimer, 2016: 316.

Gymnognathus moranus Jordan, 1937: 222

Distribution. Costa Rica [type].

Gymnognathus nanus Jordan, 1904: 248

Distribution. Brazil [type].

Gymnognathus nebulosus Motschulsky, 1875: 235

Distribution. Brazil [type].

Gymnognathus nica Jordan, 1937: 222

Distribution. Brazil [type].

Gymnognathus nubilus Jordan, 1904: 246

Distribution. Brazil [type].

Gymnognathus ophiopis Dalman, 1833: 163

Distribution. Nicaragua, Panama, Colombia, Brazil.

Gymnognathus ophiopsis Gymnognathus ophiopsis
Gymnognathus ophiopsis Dalman: 1.) lateral view, 2.) dorsal view. Specimen deposited at Arizona State University Biocollections (ASUCOB, ASUHIC). Images and data available at ecdysis portal and GBIF. Images: Samanta Orellana.

Gymnognathus ornatus Jordan, 1895: 171

Distribution. Brazil [type].

Gymnognathus pentilus Jordan, 1937: 225

Distribution. French Guiana [type].

Images. Rheinheimer, 2016: 315.

Gymnognathus phanerus Jordan, 1937: 229

Distribution. Brazil, Bolivia, French Guiana.

Images. Rheinheimer, 2016: 316.

Gymnognathus phanerus Gymnognathus phanerus
Gymnognathus phanerus Jordan, 1.) lateral view, 2.) dorsal view. Specimens deposited at Kansas University Natural History Museum (Curator: Dr. Zachary Falin), on loan at State University Biocollections (2022-2024). Images: Samanta Orellana.

Gymnognathus pindonis Jordan, 1937: 231

Distribution. French Guiana.

Images. Rheinheimer, 2016: 316.

Gymnognathus polius Jordan, 1906: 337

Distribution. Panama.

Gymnognathus procerus Jordan, 1937: 226

Distribution. French Guiana.

Images. Rheinheimer, 2016: 314.

Gymnognathus pulchellus Jordan, 1937: 229

Distribution. Brazil.

Gymnognathus pulcher Jordan, 1906: 340

Distribution. Guatemala, Panama.

Gymnognathus regalis Jordan, 1937: 225

Distribution. Brazil, French Guiana.

Images. Rheinheimer, 2016: 315.

Gymnognathus robustus Jordan, 1895: 169

Distribution. Brazil.

Images. Rheinheimer, 2004: 240.

Gymnognathus robustus Gymnognathus robustus
Gymnognathus robustus Jordan: 1.) lateral view, 2.) dorsal view. Specimen deposited at Arizona State University Biocollections (ASUCOB, ASUHIC). Images and data available at ecdysis portal and GBIF. Images: Samanta Orellana.

Gymnognathus ruficlava Jordan, 1906: 339

Distribution. Panama.

Gymnognathus ruficlava
Gymnognathus ruficlava Jordan: 1.) dorsal view. Illustration in Jordan 1906, digitized by the Smithsonian Libraries under a CC0-Public Domain License. Modified by Samanta Orellana.

Gymnognathus scalaris Jordan, 1906: 336

Distribution. Mexico, Nicaragua.

Gymnognathus scolytinus Jordan, 1904: 252

Distribution. Peru, French Guiana.

Images. Rheinheimer, 2016: 313.

Gymnognathus signatus Gyllenhal, 1833: 165

Distribution. Brazil.

Gymnognathus soror Jordan, 1904: 249

Distribution. Brazil.

Gymnognathus stigmenus Wolfrum, 1955: 676

Distribution. Brazil.

Gymnognathus talis Jordan, 1937: 230

Distribution. French Guiana.

Images. Rheinheimer, 2016: 317.

Gymnognathus tenuis Jordan, 1895: 250

Distribution. Brazil.

Gymnognathus thecla Jordan, 1906: 338

Distribution. Panama.

Gymnognathus triangularis Valentine, 1998: 269

Distribution. United States of America.

Gymnognathus uta Wolfrum, 1955: 676

Distribution. Brazil.

Gymnognathus vanda Jordan, 1937: 219

Distribution. Brazil, French Guiana.

Gymnognathus variicornis Jordan, 1895: 174

Distribution. Brazil.

Gymnognathus vicinus Jekel, 1855: 117

Distribution. Brazil.

Images. Rheinheimer, 2016: 316.

Gymnognathus viticollis Jordan, 1897: 175

Distribution. Bolivia.

Gymnognathus groups according to Jordan (1937)

SectionSubsectionGroupSpeciesDistributionImage
Section I (Pronotum without longitudinal stripes)Subsection A (Head and apex of pronotum red)Group a (Pygidium brown with white borders)G. bolshi Jordan, 1985ParaguayEugonodes brevirostris
   G. cincticollis Jordan, 1895BrazilEugonus particolor
   G. robustus Jordan, 1895BrazilEugonodes brevirostris
   G. hetarus Jordan, 1937Brazil, French GuianaEugonodes brevirostris
   G. emma Jordan, 1904BrazilEugonodes brevirostris
   G. calus Jordan, 1906PanamaG calus
   G. pulcher Jordan, 1906GuatemalaEugonodes brevirostris
   G. helma Jordan, 1937BrazilEugonodes brevirostris
  Group b (Pygidium brown with mesial stripe)G. ornatus Jordan, 1895BrazilEugonodes brevirostris
   G. scolytinus Jordan, 1904PeruEugonodes brevirostris
   G. comptus Jordan, 1904BrazilG comptus
   G. clathratus Jordan, 1895BrazilEugonodes brevirostris
   G. coronatus Jordan, 1904Panama, BrazilEugonodes brevirostris
   G. bryanti Jordan, 1937BrazilEugonodes brevirostris
   G. ruficlava Jordan, 1906 G ruficlava
   G. indagatus Wolfrum, 1931Costa RicaEugonodes brevirostris
 Subsection B (Occipucium with orange pubescence)Group cG. bella Jordan, 1904BrazilEugonodes brevirostris
   G. clara Jordan, 1904BrazilEugonodes brevirostris

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